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Laura -The nature of culture

I am kind of confused by this article but also intrigued. Grosz talks about three characteristics of nature: the force of development, the force of variation and the force of differences in sex and race. She then goes on to state that "Culture can be regarded as the varying innovative responses to the problems that nature poses to the living". Doesn't this imply that nature must come first in order to have cultural responses to it? Nature is then the force or framework of culture.
However at the end of the article she poses questions that seem to have been already answered. She asks "What would the study of culture, cultural studies, look like if nature was regarded as the framework and provocation of culture rather than its retardation?".